The OP is reporting that when he runs his the bolt out of the project rifle in the new rifles, it works fine, despite the galling on the lugs.

Conversely when he runs his new bolts in the donor rifle, they do not work fine despite a new clean bolt lug surface.

What they have in common is the bolt lug cuts in the receiver.

I would do a close examination of the receiver bolt lug cutouts, that would seem to be damaged.