Only difference is the stock. Which I would recommend really depends on how you intend to use it and what kind of stock you prefer.
If you want a walking/stalking hunting rifle, go with the Walking Varminter. The stock isn't the best, but it's sufficient and it's light whereas the factory laminate stocks are heavy and have a wide forearm that doesn't fit the hand as well for off-hand shots. It's not bad off-hand, but not ideal IMO. If you can't stand the stock Boyds offers some sporter-style replacements in walnut and laminate (Prairie Hunter, Classic & Heritage). They'll ad a little weight but still be comfortable for off-hand work.
If you shoot mostly from a bench or bipod, go with the Lightweight Varminter with factory laminate stock. Personally I hate thumbhole stocks, but a lot of guys swear by them so choose whichever tickles your fancy.
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